Key combinations for startup options not working

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Here's a puzzle for the Mac experts.
None of the key combinations to access the startup options, including safe mode, recovery, single user mode, startup disk etc on my Mac Pro mid 2012 5,1 have any effect. I'm using OS 10.14.6 with a standard Apple wired keyboard connected to one of the USB 2 ports at the front of the Mac Pro.
Resetting the NVRAM using alt+cmd+p+r gave the repeated bongs but didn't fix this. No other key combos work. Swapping the keyboard doesn't help.
I had this same problem on my previous Mac Pro so it seems that I must have imported the cause when I migrated the contents of the old Mac to this Mac.
What could be causing this?
David Hoffman
 

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